What is moral? I know we have what have been called "mirror neurons" to aide in empathy. For example when I get punched in the shoulder it may actually hurt me, but when someone witnesses the visual of it they can practically feel the impact themselves with a neuron located near the actual sensory neuron in the viewer. But is this true empathy? When apes see someone in their group choking they all as a group start coughing in the event that the choking one ate poison that they have all possibly consumed. This is just a group survival behavior. It does nothing towards empathy towards other forms of life that ate not as familiar.
On a different note a person who is a psychopath may know the harm they do, but they don't empathize about it in the sense of feeling it. Are they immoral due to biological deficiency? Their intelligence is certainly there.